) Redhead et al., and L. velutina (Quél.) Redhead et al. Species included based on morphology (Redhead et al. 2002) are L. aurantiaca (Redhead & Kuyper) Redhead et al., L. chromacea

(Cleland) Redhead et al., and L. lobata (Redhead & Kuyper) Redhead et al. Comments Subg. Lichenomphalia forms a Anlotinib mouse well-supported, monophyletic clade that is concordant with the morphological and ecological characters that define the group. Species in subg. Lichenomphalia are found in high-light habitats that are more subject to drought than in subg. Protolichenomphalia, but they are presumably protected from ionizing radiation and desiccation by strong pigments and thick hyphal walls in the thalli (Redhead et al. 2002; Redhead and Kuyper 1987). Lichenomphalia subgen. Protolichenomphalia Lücking, Redhead & Novell, subg. nov. Mycobank MB 804123. Type species: Lichenomphalia umbellifera (L.) Redhead, A-1210477 manufacturer Lutzoni, Moncalvo & Vilgalys, Mycotaxon 83: 38 (2002) ≡ Agaricus umbelliferus L., Sp. pl. 2: 1175 (1753), sanctioned by Fr., Elench. fung. 1: 22 (1828). Etymology—proto – first, lichenomphalia – Lichenomphalia. IWR1 Characters as in Lichenomphalia, basidiomes lightly pigmented; lichenized thallus undifferentiated, hyphal walls thin; growing in mesic habitats in arctic and boreal zones. Phylogenetic support Phylogenetic

support is irrelevant as this subgenus is monotypic. Species included Type species: Lichenomphalia umbellifera. Comments Redhead et al. (2002) noted that L. umbellifera has more ancestral features than other species now placed in subg. Lichenomphalia, i.e., Protein tyrosine phosphatase the hyphae in the thallus are broader and not as thick-walled, so presumably more susceptible to desiccation (Redhead and Kuyper 1988). Furthermore, the type of subg. Protolichenomphalia has a broader geographical distribution, occupies wetter habitats, and its basidiomata are less protected by strong pigments than species in subg. Lichenomphalia (Redhead et al. 2002; Lawrey et al. 2009). Semiomphalina Redhead, Can. J.

Bot. 62(5): 886 (1984). Type species: Semiomphalina leptoglossoides (Corner) Redhead, Can. J. Bot. 62(5): 886 (1984), ≡ Pseudocraterellus leptoglossoides Corner, Monogr. Cantharelloid Fungi: 161 (1966). Basidiomes arrhenioid, drooping, pale; stipe and thallus similar to those of Lichenomphalia umbellifera. Comments There are currently no published sequences of this lichenized, monotypic genus described from Papua New Guinea by Corner, but Redhead et al. (2002) suggested that it was related to Lichenomphalia based on morphology and ecology. If Semiomphalina leptoglossoides and Lichenomphalia hudsoniana are later found to be congeneric, Article 14 in the Melbourne Code (2012) allows for selection of a widely used name, such as Lichenomphalia, over a more obscure one (Semiomphalina). Tribe Cantharelluleae Lodge, Redhead, Norvell & Desjardin, tribe nov. MycoBank MB804125. Type genus: Cantharellula Singer, Revue Mycol., Paris 1: 281 (1936).